Epithelia are multiscale active liquid crystals

JM Armengol-Collado, LN Carenza, J Eckert… - Nature Physics, 2023 - nature.com
Biological processes such as embryogenesis, wound healing and cancer progression rely
on the ability of epithelial cells to coordinate their mechanical activity over length scales that …

A geometric-tension-dynamics model of epithelial convergent extension

NH Claussen, F Brauns, BI Shraiman - Proceedings of the National …, 2024 - pnas.org
Convergent extension of epithelial tissue is a key motif of animal morphogenesis. On a
coarse scale, cell motion resembles laminar fluid flow; yet in contrast to a fluid, epithelial …

Hexanematic crossover in epithelial monolayers depends on cell adhesion and cell density

J Eckert, B Ladoux, RM Mège, L Giomi… - Nature …, 2023 - nature.com
Abstract Changes in tissue geometry during developmental processes are associated with
collective migration of cells. Recent experimental and numerical results suggest that these …

Hydrodynamic theory of -atic liquid crystals

L Giomi, J Toner, N Sarkar - Physical Review E, 2022 - APS
We formulate a comprehensive hydrodynamic theory of two-dimensional liquid crystals with
generic p-fold rotational symmetry, also known as p-atics, of which nematics (p= 2) and …

Cell Division and Motility Enable Hexatic Order in Biological Tissues

Y Tang, S Chen, MJ Bowick, D Bi - Physical Review Letters, 2024 - APS
Biological tissues transform between solid-and liquidlike states in many fundamental
physiological events. Recent experimental observations further suggest that in two …

Long-range order in two-dimensional systems with fluctuating active stresses

YE Keta, S Henkes - ar** germ layers, pair-wise forces (or active
stresses) arise from the contractile activity of the cytoskeleton, with dissipation provided by …

Alignment rule and geometric confinement lead to stability of a vortex in active flow

YH Zhang, Z Yao - The European Physical Journal E, 2023 - Springer
Vortices are hallmarks of a wide range of nonequilibrium phenomena in fluids at multiple
length scales. In this work, we numerically study the whirling motion of self-propelled soft …

Coarse-graining dense, deformable active particles

MR Nejad, JM Yeomans - arxiv preprint arxiv:2501.07280, 2025 - arxiv.org
We coarse-grain a model of closely-packed ellipses that can vary their aspect ratio to derive
continuum equations for materials comprising confluent deformable particles such as …

Controlling flow patterns and topology in active emulsions

G Negro, LC Head, LN Carenza, TN Shendruk… - arxiv preprint arxiv …, 2024 - arxiv.org
Active emulsions and liquid crystalline shells are intriguing and experimentally realisable
types of topological matter. Here we numerically study the morphology and spatiotemporal …

Phase diagram, confining strings, and a new universality class in nematopolar matter

F Vafa, A Doostmohammadi - arxiv preprint arxiv:2501.04769, 2025 - arxiv.org
We study a minimal model of a system with coexisting nematic and polar orientational
orders, where one field tends to order and the other prefers isotropy. For strong coupling, the …